Cortinarius helodes M. M. Moser, Matheny & Daniele + Alnus

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Literature

• references
  Becerra A, Nouhra E, Daniele G, Domínguez L, McKay D (2005) Ectomycorrhizas of Cortinarius helodes and Gyrodon monticola with Alnus acuminata from Argentina. Mycorrhiza 15(1): 7-15.

Morphology

Morphology of the mycorrhizal system
• ramification presence-type
  absent
• rhizomorphs as stout, short, conical structures presence-abundance
  absent
• rhizomorphs presence
  present
• exploration type
  medium distance fringe
Morphology of the unramified ends
• shape
  straight
or    bent
• shape {of distal end}
  not inflated, cylindric
or    tapering
• length
1.2 mm    Lower value of unspecified range (could be µ-s.d., but not known)
9.2 mm    Upper value of unspecified range (could be µ+s.d., but not known)
• diameter
0.4 mm    Lower value of unspecified range (could be µ-s.d., but not known)
0.8 mm    Upper value of unspecified range (could be µ+s.d., but not known)
• colour
  white
• very tip colour
  lilac, light reddish blue [with bluish to lilac tints towards the apex]
• older parts colour
  ochre, yellowish brown
or    yellowish
• mantle laticifers visibility
  absent
• mantle secreted latex {after scratching mantle} presence
  present [young mycorrhizae exude whitish substace when damaged]
• mantle secreted latex {after scratching mantle} colour-habit
  white, remaining white after some minutes
• mantle carbonizing presence
  absent
• mantle surface {in general} habit
  smooth
or    covered with soil particles
or    not smooth
• mantle surface {in detail} kind
  loosely woolly
• emanating hyphae presence
  present
• emanating hyphae abundance
  infrequent
or    abundant
Morphology of rhizomorphs
• diameter
0.04 mm    Mean (= average)
• colour
  yellowish
or    white
Morphology of sclerotia
• presence
  absent

Anatomy

Anatomical features of the entire mycorrhiza
• emanating elements presence-type
  rhizomorphs
or    emanating hyphae
• crystals presence
  absent
• blue granules presence
  absent
Anatomy of laticifers
• presence
  absent
• anatomy entire mycorrhiza laticiferous solitary (gloeoplerous) cells presence
  absent
Anatomical features of the mantle
• completeness
  incomplete
or    complete
Anatomy of the outer mantle layer apart from the ectormycorrhizal tip
• organisation
  plectenchymatous
• mantle type
  hyphae rather irregularly arranged and no special pattern discernible (type B)
• hyphal system kind
  undifferentiated
• hyphae arrangement
  not specific [hyphae tangentially oriented]
• septa clamps presence
  present
• cell pigment location-colour
  absent
• cell diameter
2.4 µm    Lower value of unspecified range (could be µ-s.d., but not known)
5 µm    Upper value of unspecified range (could be µ+s.d., but not known)
• cell wall thickness
0.1 µm    Mean (= average)
• cell wall surface habit
  smooth
• roundish plasmatically yellow cells presence
  absent
• pigment granula on surface presence
  absent
Anatomy of the middle mantle layer
• presence
  present
• organisation
  pseudoparenchymatous
• hyphae arrangement
  plectenchymatous, without pattern
• cell shape
  polygonal
or    roundish
• cell pigment location-colour
  colourless
• cell diameter
4 µm    Lower value of unspecified range (could be µ-s.d., but not known)
24 µm    Upper value of unspecified range (could be µ+s.d., but not known)
• cell length
6 µm    Lower value of unspecified range (could be µ-s.d., but not known)
24 µm    Upper value of unspecified range (could be µ+s.d., but not known)
Anatomy of the inner mantle layer
• organisation
  pseudoparenchymatous
• cell shape
  polygonal
or    roundish
• hyphae vessel-like presence
  absent
• cell pigment location-colour
  absent
• cell diameter
3.2 µm    Lower value of unspecified range (could be µ-s.d., but not known)
11 µm    Upper value of unspecified range (could be µ+s.d., but not known)
• cell length
3.2 µm    Lower value of unspecified range (could be µ-s.d., but not known)
11 µm    Upper value of unspecified range (could be µ+s.d., but not known)
Anatomy of the mantle in longitudinal section
• mantle thickness {apart from tip}
130 µm    Lower value of unspecified range (could be µ-s.d., but not known)
220 µm    Upper value of unspecified range (could be µ+s.d., but not known)



Anatomy of the mantle in cross-section
• mantle different layers presence
  discernible
• outer mantle layer organisation
  plectenchymatous
• outer mantle layer hyphae radially diameter
3.2 µm    Lower value of unspecified range (could be µ-s.d., but not known)
6.5 µm    Upper value of unspecified range (could be µ+s.d., but not known)
• middle mantle layer presence
  present
• middle mantle layer organisation
  pseudoparenchymatous
• middle mantle layer hyphae tangentially length
6.5 µm    Lower value of unspecified range (could be µ-s.d., but not known)
27.5 µm    Upper value of unspecified range (could be µ+s.d., but not known)
• middle mantle layer hyphae radially diameter
6.5 µm    Lower value of unspecified range (could be µ-s.d., but not known)
27.5 µm    Upper value of unspecified range (could be µ+s.d., but not known)
• inner mantle layer organisation
  pseudoparenchymatous
• inner mantle layer hyphae tangentially length
4 µm    Lower value of unspecified range (could be µ-s.d., but not known)
16 µm    Upper value of unspecified range (could be µ+s.d., but not known)
• inner mantle layer hyphae radially diameter
4 µm    Lower value of unspecified range (could be µ-s.d., but not known)
16 µm    Upper value of unspecified range (could be µ+s.d., but not known)

Anatomy of the tannin cells in cross-section
• presence
  absent

Anatomy of the cortical cells in cross-section
• anatomy mantle cross-section cortical (epidermal) cells shape
  tangentially-oval to tangentially-elliptic
• anatomy mantle cross-section cortical (epidermal) cells tangentially length
8 µm    Lower value of unspecified range (could be µ-s.d., but not known)
30.5 µm    Upper value of unspecified range (could be µ+s.d., but not known)
• anatomy mantle cross-section cortical (epidermal) cells radially diameter
6.5 µm    Lower value of unspecified range (could be µ-s.d., but not known)
19.5 µm    Upper value of unspecified range (could be µ+s.d., but not known)

Anatomy of the Hartig net in cross-section
• presence
  present
• kind
  paraepidermal
• anatomy mantle cross-section hyphal cells around cortical (epidermal) cells shape
  roundish
or    cylindrical
• anatomy mantle cross-section hyphal cells around cortical (epidermal) cells thickness
1.6 µm    Lower value of unspecified range (could be µ-s.d., but not known)
5 µm    Upper value of unspecified range (could be µ+s.d., but not known)
• anatomy mantle cross-section hyphal rows around cortical (epidermal) cells number
  one
Anatomy of the emanating elements in general
• anastomoses type
  open, with a short bridge or bridge almost lacking
Anatomy of cystidia


Anatomy of emanating hyphae
• shape
  straight
or    not striking
• cell pigment location-colour
  absent

Anatomy of the ramification of emanating hyphae
• presence
  present

Anatomy of the septa of emanating hyphae
• clamps presence
  present

Anatomy of secondary septa of emanating hyphae
• presence
  present
• abundance
  abundant

Anatomy of the cells of emanating hyphae
• anatomy emanating elements emanating hyphae cell diameter
2.5 µm    Lower value of unspecified range (could be µ-s.d., but not known)
6.5 µm    Upper value of unspecified range (could be µ+s.d., but not known)
• anatomy emanating elements emanating hyphae cell wall surface habit
  smooth
or    with soil particles


Anatomy of the warts occouring on emanating hyphae
• presence
  absent
Anatomy of rhizomorphs
• type
  undifferentiated; hyphae rather loosely woven and of uniform diameter
(type A)
• hyphae distinctly different diameter
  absent

Anatomy of hyphae in rhizomorphs
• anatomy emanating elements rhizomorphs clamps presence
  present, not specified
Anatomy of vessel-like or thickened hyphae in rhizomorphs
• anatomy emanating elements rhizomorphs clamps presence
  present, not specified
Anatomy of central non-vessel-like hyphae in rhizomorphs
• anatomy emanating elements rhizomorphs clamps presence
  present, not specified
Anatomy of peripheral hyphae in rhizomorphs
• anatomy emanating elements rhizomorphs clamps presence
  present, not specified
Anatomy of solitary thick-walled hyphae in rhizomorphs
• anatomy emanating elements rhizomorphs clamps presence
  present, not specified
Anatomy of chlamydospores
• presence
  absent
Anatomy of haustoria
• {of ectomycorrhiza former} presence
  present
• shape
  as normal hyphae

Chemical reactions

• reaction with cotton-blue-lactic-acid presence
  present [slightly blue]
• reaction with ethanol 70% presence
  absent
• reaction with KOH 10% presence
  absent
• reaction with lactic acid presence
  absent
• reaction with Melzer's reagent presence
  absent
• reaction with sulpho-vanillin presence
  absent

Ecology

• geographic occurrence continent
  South America

Tree

• plant family
  Betulaceae
• plant genus
  Alnus
• plant habitat kind
  forests, woods

Fungus

• family
  Cortinariaceae
• fruitbodies growth habit
  epigeous
or    pileate-lamellate

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